1. Department of Biological Sciences, Towson University, Towson, Maryland 21252;
2. Department of Evolution, Biological Faculty, M.V. Lomonosov Moscow State University, Vorobjovy gory, 119992, Moscow, Russia;
3. Laboratoire Géosciences & Observatoire des Sciences de l'Univers de Rennes, Université Rennes 1, 35042 Rennes, France;