1. Wellcome Sanger Institute, Wellcome Genome Campus, Hinxton CB10 1SA, United Kingdom;
2. European Molecular Biology Laboratory (EMBL)–European Bioinformatics Institute (EBI), Wellcome Genome Campus, Hinxton CB10 1SD, United Kingdom
3. Theory of Condensed Matter Research Group, Cavendish Laboratory, University of Cambridge, Cambridge CB3 0HE, United Kingdom