1. The Francis Crick Institute, London NW1 1AT, United Kingdom;
2. Department of Genetics, Environment and Evolution, UCL Genetics Institute, University College London, London WC1E 6BT, United Kingdom
3. Department of Molecular Neuroscience, UCL Institute of Neurology, University College London, London WC1E 6BT, United Kingdom
4. Okinawa Institute of Science and Technology Graduate University, Onna-son, Okinawa 904-0412, Japan