1. Department of Agroecology, Aarhus University, DK-4200, Slagelse, Denmark;
2. Rothamsted Research, Harpenden, AL5 2JQ, United Kingdom
3. Centre for Crop and Disease Management (CCDM), Curtin University, Bentley, Western Australia, WA-6102, Australia
4. ADAS High Mowthorpe, Malton, North Yorkshire, YO17 8BP, United Kingdom