1. Department of Biological Sciences, University of Tasmania, Hobart, Tasmania, Australia; email:
2. Odum School of Ecology, University of Georgia, Athens, Georgia, USA; email:
3. Department of Archaeology and History, La Trobe University, Bundoora, Victoria, Australia; email:
4. Caesar Kleberg Wildlife Research Institute, Texas A&M University–Kingsville, Kingsville, Texas, USA; email: