1. Biology Department, College of William and Mary, Williamsburg, Virginia 23185;
2. Department of Plant and Soil Sciences, University of Delaware, Newark, Delaware 19716
3. Department Biological Sciences, University of Delaware, Newark, Delaware 19716
4. College of Earth, Ocean, and Environment, University of Delaware, Newark, Delaware 19716
5. Biosystems Engineering and Soil Science Department, University of Tennessee, Knoxville, Tennessee 37996