1. Department of Biology, Washington University, St. Louis, Missouri 63130, USA
2. Department of Biochemistry and Molecular Biophysics, Washington University School of Medicine, St. Louis, Missouri 63110, USA;
3. Center for Science and Engineering of Living Systems, Washington University, St. Louis, Missouri 63130, USA
4. Center for Engineering MechanoBiology, Washington University, St. Louis, Missouri 63130, USA
5. Department of Biology, Duke University, Durham, North Carolina 27708, USA;