1. Department of Plant Biology, University of Illinois at Urbana-Champaign, Urbana-Champaign, Illinois 61801;
2. INRA-CNRS, UMR1165, Unité de Recherche en Génomique Végétale, F-91057 Evry, France;
3. Department of Plant Production, College of Food and Agricultural Sciences, King Saud University, Riyadh 11451, Saudi Arabia
4. Department of Biology, University of Munich, D-80638 Munich, Germany;