1. Jealott's Hill International Research Centre, Syngenta, Bracknell RG42 6EY, United Kingdom;
2. Institute of Plant and Microbial Biology, Academia Sinica, Taipei 11529, Taiwan;
3. The Sainsbury Laboratory, University of East Anglia, Norwich NR4 7UH, United Kingdom
4. Current affiliation: Institute of Biology Leiden, Leiden University, Leiden 2333 BE, The Netherlands;