1. Department of Earth and Environment, Boston University, Boston, Massachusetts 02215;,
2. Division of Biological Sciences, University of Montana, Missoula, Montana 59812;
3. Earth Systems Research Center, University of New Hampshire, Durham, New Hampshire 03824;
4. Department of Earth System Science, University of California, Irvine, California 92697-3100;
5. Department of Organismic and Evolutionary Biology, Harvard University, Cambridge, Massachusetts 02138;
6. Department of Plant and Soil Sciences and Delaware Environmental Institute, University of Delaware, Newark, Delaware 19716;