1. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China;
2. Department of Ocean Science, Hong Kong University of Science and Technology, Hong Kong, China
3. State Key Laboratory of Marine Environmental Science, College of Ocean and Earth Sciences, Institute of Marine Microbes and Ecospheres, Xiamen University (Xiang'an), Xiamen, Fujian, China
4. Shenzhen Key Laboratory of Marine Bioresource and Eco-Environmental Science, College of Life Sciences and Oceanography, Shenzhen University, Shenzhen, China;
5. Institute for Advanced Study, Shenzhen University, Shenzhen, China;