1. Natural Resources Science and Management, University of Minnesota, St. Paul, Minnesota, 55108, USA;
2. Department of Bioproducts and Biosystems Engineering, University of Minnesota, St. Paul, Minnesota, 55108, USA;
3. Department of Ecology, Evolution, and Behavior, University of Minnesota, St. Paul, Minnesota, 55108, USA;
4. Bren School of Environmental Science and Management, University of California, Santa Barbara, California, 93117, USA