1. Dauphin Island Sea Lab, Dauphin Island, Alabama 36528;
2. National Oceanography Centre, Southampton, School of Ocean and Earth Science, University of Southampton, Southampton SO14 3ZH, United Kingdom;
3. British Antarctic Survey, NERC, Cambridge CB3 0ET, United Kingdom;,
4. Department of Geology, University of Illinois, Urbana, Illinois 61801;
5. Department of Biological Sciences, University of Rhode Island, Kingston, Rhode Island 02881;,