1. Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut 06269;email:
2. School of Biological Sciences, Victoria University of Wellington, Wellington 6014, New Zealand
3. Landcare Research, Auckland 1142, New Zealand;
4. Department of Evolutionary Biology, University of Siena, 53100 Siena, Italy;
5. Department of Molecular Biology and Biochemistry, Simon Fraser University, Burnaby, British Columbia V5A 1S6, Canada;,
6. Department of Laboratory Medicine, Division of Metabolic Diseases, Karolinska Institutet, Norvum 141 86, Stockholm, Sweden