1. Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey 08544;
2. National Center for Ecological Analysis and Synthesis, 735 State Street, Santa Barbara, California 93101;
3. Department of Life Sciences, Ben-Gurion University of the Negev, Beer-Sheva 84105, Israel;
4. Evolution et Diversité Biologique, CNRS, Université Paul Sabatier, bâtiment IVR3, F-31062 Toulouse, France;