1. Department of Biology, Faculty of Science, 2Research and Development Center for Higher Education, Kyushu University, Ropponmatsu, Fukuoka, 810-8560, Japan;,
2. Department of Biology, Pennsylvania State University, University Park, Pennsylvania 16802-5301;
3. PRESTO, Japan Science and Technology Agency, 4-1-8 Honcho Kawaguchi, Saitama, Japan;