1. School of Biological Sciences, University of Exeter, Hatherly Laboratories, Prince of Wales Road, Exeter, EX4 4PS, United Kingdom;
2. Boyce Thompson Institute for Plant Research at Cornell University, Tower Road, Ithaca, NY 14853;
3. Institute of Biological Chemistry, Washington State University, P.O. Box 646340, Pullman, WA 99164-6340;