1. Department of Biology, Lund University, Sölvegatan 37, SE-223 62 Lund, Sweden;
2. Universidade de Lisboa, Faculdade de Ciências, Centro de Oceanografia, Campo Grande, 1749-016 Lisboa, Portugal
3. Institut des Sciences de l'Evolution, UMR 5554 CNRS/UM2/IRD, Université Montpellier 2, cc065, place Eugène Bataillon, 34095 Montpellier cedex 05, France