1. Department of Biology, Faculty of Sciences, Ferdowsi University of Mashhad, Mashhad, Iran;
2. Rodentology Research Department, Ferdowsi University of Mashhad, Mashhad, Iran
3. Zoological Innovations Research Department, Institute of Applied Zoology, Faculty of Sciences, Ferdowsi University of Mashhad, Mashhad, Iran
4. Zoological Institute, Russian Academy of Sciences, St-Petersburg 199034, Russia
5. Slovenian Museum of Natural History, Prešernova 20, SI-1000 Ljubljana, Slovenia