1. Hartpury University, Gloucestershire, United Kingdom; e-mail: lucy.bearman-brown@hartpury.ac.uk
2. Conservation K9 Consultancy, Wrexham, United Kingdom; e-mail: louise@conservationk9consultancy.com
3. School of Biological Sciences, University of Reading, Reading, United Kingdom; e-mail: p.j.baker@reading.ac.uk, L.C.Evans@pgr.reading.ac.uk