1. Mammal Research Institute, Faculty of Natural and Agricultural Sciences, University of Pretoria, Hatfield, South Africa; e-mail: mark.keith@up.ac.za, mariette.e.pretorius@gmail.com
2. Department of Medical Virology, Faculty of Health Sciences, Centre for Viral Zoonoses, University of Pretoria, South Africa; e-mail: wanda.markotter@up.ac.za
3. African Bats NPC, South Africa; e-mail: ernest.seamark@africanbats.org
4. Department of Biology, University of Waterloo, Waterloo, Ontario, Canada; e-mail: hugh.broders@uwaterloo.ca