1. Natural History Museum, London, UK; e-mail:
2. Department of Ecology and Evolutionary Biology, University of Arizona, Tucson, USA; e-mail:
3. Aquatic and Terrestrial Ecology, Royal Belgian Institute of Natural Science, Brussel, Belgium; e-mail:
4. Laboratorio de Sistemática Molecular, Carrera de Biología, Universidad Nacional Autónoma de México, Col. Ejército de Oriente, Ciudad de México, México; e-mail:
5. Amphibian and Reptile Diversity Research Center, Department of Biology, University of Texas at Arlington, Arlington, USA; e-mail: