1. Roztocze National Park, Zwierzyniec, Poland; e-mail:
2. Department of Anatomy and Comparative Anthropology, Institute of Biology, University of Maria Sklodowska-Curie, Lublin, Poland; e-mail:
3. Department of Ecology, Institute of Functional Biology and Ecology, Faculty of Biology, University of Warsaw, Biological and Chemical Research Centre, Warszawa, Poland; e-mail: