1. School of Natural Resources, University of Missouri, Columbia, Missouri, U.S.A.
2. Department of Wildlife, Fisheries, and Conservation Biology, University of Maine, Orono, Maine, U.S.A.
3. Caesar Kleberg Wildlife Research Institute, Texas A&M University-Kingsville, Kingsville, Texas, U.S.A.
4. Department of Biology, University of Saskatchewan, Saskatoon, Saskatchewan, Canada