1. Department of Biology, Virginia Commonwealth University, 1000 West Cary Street, Richmond, Virginia, 23284, USA
2. School of Environmental and Natural Resources, The Ohio State University, 2021 Coffey Road, Columbus, Ohio, 43210, USA
3. Center for Environmental Studies, Virginia Commonwealth University, 1000 West Cary Street, Richmond, Virginia, 23284, USA
4. Migratory Bird Center, Smithsonian Conservation Biology Institute, P.O. Box 37012 MRC 5503, Washington, D.C., 20013, USA