1. U.S. Geological Survey, Eastern Ecological Science Center at Patuxent Wildlife Refuge, 12100 Beach Forest Road, Laurel, Maryland, 20708, USA
2. Department of Zoology, Southern Illinois University, Life Science II, Carbondale, IL 62901, USA
3. Department of Natural Resources, Maryland Natural Heritage Program, Annapolis, MD 21401, USA.
4. U.S. Fish and Wildlife Service, Chesapeake Bay Field Office, 177 Admiral Cochran Drive, Annapolis, Maryland, 21401, USA
5. No Affiliation