1. College of Nature Conservation, Beijing Forestry University, No.35 East Qinghua Road, Haidian, Beijing, 100083, China
2. Gansu Yanchiwan National Natural Reserve Administration Bureau, No.12 Bayin Road, Subei, Gansu, 736300, China
3. Beijing Wildlife Ark, No.34-01-801 Tiantongyuan Community, Changping, Beijing, 102218, China