1. Department of Forestry and Environmental Conservation and South Carolina Cooperative Fish and Wildlife Research Unit, Clemson University, Clemson, South Carolina, 29634, USA
2. Department of Natural Resources Science, University of Rhode Island, Kingston, Rhode Island, 02881, USA
3. Dauphin Island Sea Lab, Dauphin Island, Alabama, 36528, USA
4. U.S. Department of the Interior, Geological Survey South Carolina Cooperative Fish and Wildlife Research Unit, Clemson University, Clemson, South Carolina, 29634, USA