1. 1Department of Biological Sciences, Western Michigan University, Kalamazoo, Michigan 49008-5200, USA
2. 2Department of Forest Biological Sciences, College of Forestry and Natural Resources, University of the Philippines Los Baños, Laguna, Philippines
3. 4Graduate School, University of the Philippines Los Baños, Laguna, Philippines
4. 5College of Sciences, Palawan State University, Tiniguiban Heights, Puerto Princesa City, Palawan, 5300 Philippines
5. 6Philippine Taxonomic Initiative, Inc., Botanica Building, El Nido, Palawan, 5313 Philippines