1. 1Shirley C. Tucker Herbarium, Department of Biological Sciences, Louisiana State University, Baton Rouge, Louisiana 70803-1705, USA; ; daniel.santamaria366@gmail.com; llagomarsino1@lsu.edu, Email: santamaria1@lsu.edu
2. 3Los Charcos de Osa Centro de Diversidad de Plantas Regionales Osa, Puntarenas, Costa Rica;, Email: reinaldo.aguilar.f@gmail.com