1. Joint Carnegie Mellon–University of Pittsburgh Ph.D. Program in Computational Biology, Department of Computation and Systems Biology, University of Pittsburgh, Pittsburgh, PA 15260, USA.
2. Department of Computational and Systems Biology, University of Pittsburgh School of Medicine, Pittsburgh, PA 15260, USA.
3. Department of Developmental Biology, University of Pittsburgh School of Medicine, Pittsburgh, PA 15201, USA.
4. University of Pittsburgh School of Medicine, Pittsburgh, PA 15261, USA.