1. Laboratory of Host Defense, WPI Immunology Frontier Research Center, Osaka University, Suita, Osaka 565–0871, Japan.
2. Department of Respiratory Medicine, Tohoku University Graduate School of Medicine, Sendai, Miyagi 980–8575, Japan.
3. Department of Molecular Protozoology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka 565–0871, Japan.
4. The Research Foundation for Microbial Diseases of Osaka University, Kan-on-ji, Kagawa 768–0061, Japan.
5. Department of Chemical Processes and Environments, University of Kitakyushu, Kitakyushu, Fukuoka 808–0135, Japan.