1. Wildlife Biology Program, W.A. Franke College of Forestry and Conservation, University of Montana, Missoula, MT, USA.
2. Montana Fish, Wildlife & Parks, Missoula, MT, USA.
3. Northern Rocky Mountain Science Center, U.S. Geological Survey, West Glacier, MT, USA.
4. Flathead Biological Station, University of Montana, Polson, MT, USA.
5. Northern Rocky Mountain Science Center, U.S. Geological Survey, Bozeman, MT, USA.