1. Epigenetics Institute, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
2. Department of Cell and Developmental Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
3. Graduate Group in Genomics and Computational Biology, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
4. Department of Genetics, University of Pennsylvania Perelman School of Medicine, Philadelphia, PA, USA.
5. Department of Bioengineering, University of Pennsylvania, Philadelphia, PA, USA.