1. Medical Research Council Mitochondrial Biology Unit, University of Cambridge, Hills Road, CB2 0XY Cambridge, UK.
2. Division of Cell Signalling & Immunology, School of Life Sciences, University of Dundee, Dundee DD1 5EH, Scotland, UK.
3. Department of Neuroscience, Columbia University, New York, NY 10032, USA.
4. Department of Medicine, University of Cambridge, Cambridge, UK.