1. ISEM, UMR5554, Université de Montpellier, CNRS, IRD, EPHE, Place Eugène Bataillon (cc065), 34095 Montpellier cedex 05, France.
2. Behavioral Ecology and Sociobiology Unit, German Primate Center, Kellnerweg 4, 37077 Göttingen, Germany.
3. Projet Mandrillus, SODEPAL, BP 52, Bakoumba, Gabon.
4. CEFE, UMR5175, CNRS, University of Montpellier, University Paul Valery Montpellier, EPHE, 1919 route de Mende, 34293 Montpellier, France.