1. Department of Biology, School of Science, Kyushu University, Fukuoka 819-0395, Japan.
2. School of Life Sciences, Ulsan National Institute of Science and Technology, Ulsan 44919, Republic of Korea.
3. Department of Environmental Changes, Faculty of Social and Cultural Studies, Kyushu University, Fukuoka 819-0395, Japan.
4. AgroParisTech, Paris F-75231 Cedex 05, France.
5. Faculty of Arts and Science, Kyushu University, Fukuoka 819-0395, Japan.