1. Center for Cooperative Research in Biosciences (CIC bioGUNE), Basque Research and Technology Alliance (BRTA), Bizkaia Technology Park, Building 801A, 48160 Derio, Spain.
2. Neiker–Tecnalia, Berreaga Kalea 1, 48160 Derio, Spain.
3. Department of Genetics, Graduate School of Medicine, Osaka University, Japan.
4. IKERBASQUE, Basque Foundation for Science, 48013 Bilbao, Spain.