1. Department of Environmental Sciences, Wageningen University, P.O. Box 47, NL-6700 AA Wageningen, Netherlands.
2. Institut des Sciences de l’Evolution, UMR 5554, CNRS, Université de Montpellier, CC 065, Place Eugéne Bataillon, 34095 Montpellier Cedex 05, France.
3. Earth System Science Group, College of Life and Environmental Sciences, University of Exeter, Exeter EX4 4QE, UK.