1. Department of Behavioural Neurobiology, Max Planck Institute for Ornithology, Eberhard-Gwinner-Straße, 82319 Seewiesen, Germany.
2. Biosystems Science Course, Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido 0600810, Japan.
3. Department of Biology, Faculty of Science, Hokkaido University, Sapporo, Hokkaido 0600810, Japan.