1. Department of Fisheries, Wildlife, and Conservation Biology, University of Minnesota, 2003 Upper Buford Circle, St. Paul, MN 55801, USA.
2. Department of Biological Sciences, University of Manitoba, 50 Sifton Road, Winnipeg, Manitoba R3T 2N2, Canada.
3. Voyageurs National Park, 360 Highway 11 E, International Falls, MN 56649, USA.