1. Department of Marine Sciences, University of Gothenburg, Box 461, SE-450 30 Göteborg, Sweden.
2. Sven Loven Center Kristineberg, University of Gothenburg, Fiskebäckskil, Sweden.
3. Natural History Museum of Denmark, University of Copenhagen, Sølvgade 83S, 1307 Copenhagen K, Denmark.
4. Department of Biological and Environmental Sciences, University of Gothenburg, Box 461, SE-405 30 Göteborg, Sweden.