1. State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Institute of Tibetan Plateau Research, Chinese Academy of Sciences, Beijing 100101, China.
2. University of Chinese Academy of Sciences, Beijing 100049, China.
3. School of Geographical Sciences, University of Bristol, Bristol BS8 1SS, UK.
4. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, China.
5. School of Environment, Earth and Ecosystem Sciences, The Open University, Milton Keynes MK7 6AA, UK.