1. Department of Physiology, Graduate School of Medicine, Osaka Metropolitan University, Osaka 545-8585, Japan.
2. Department of Physiology, Graduate School of Medicine, Osaka City University, Osaka 545-8585, Japan.
3. Department of Life Sciences, Graduate School of Arts and Sciences, The University of Tokyo, Meguro, Tokyo 153-8902, Japan.
4. Komaba Institute for Science, The University of Tokyo, Meguro, Tokyo 153-8902, Japan.
5. PRESTO, Japan Science and Technology Agency (JST), Kawaguchi, Saitama 332-0012, Japan.