1. Division of Cell and Developmental Biology, School of Life Sciences, University of Dundee, Dundee DD1 5EH, UK.
2. Department of Physics, University of California San Diego, La Jolla, CA 92093, USA.
3. School of Engineering and Applied Sciences, Harvard University, Cambridge, MA 02134, USA.
4. Departments of Physics and Organismic and Evolutionary Biology, Harvard University, Cambridge, MA 02138, USA.