1. State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China.
2. Scripps Institution of Oceanography, University of California San Diego, La Jolla, CA 92093, USA.
3. Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering and School of Marine Science, Sun Yat-Sen University, Guangzhou, China.
4. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai), Zhuhai, China.
5. Woods Hole Oceanographic Institution, Woods Hole, MA 02543, USA.
6. Department of Earth and Planetary Sciences, University of California Riverside, Riverside, CA 92521, USA.