1. State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou, China.
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou), Guangzhou, China.
3. Innovation Academy of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences, Guangzhou, China.
4. Georgia Institute of Technology, Atlanta, GA, USA.
5. GEOMAR Helmholtz Centre for Ocean Research Kiel, Kiel, Germany.