1. Apicultural Research Institute, College of Animal Science and Technology, Yangzhou University, Yangzhou 225009, China.
2. Department of Biomedical Sciences, City University of Hong Kong, Hong Kong SAR, China.
3. Apiculture Science Institute of Jilin Province, Jilin 132108, China.
4. Mudanjiang Branch of Heilongjiang Academy of Agricultural Sciences, Mudanjiang 157043, China.