1. Department of Molecular and Cellular Biology, Howard Hughes Medical Institute, The University of California, Berkeley, Berkeley, CA 94720, USA.
2. Leonard Davis School of Gerontology, University of Southern California, Los Angeles, CA 90089.
3. Center for Bioengineering and Tissue Regeneration, Department of Surgery, University of California, San Francisco, San Francisco, CA 94143, USA.
4. Department of Genetics, Perelman School of Medicine, University of Pennsylvania, Philadephia, PA 191004, USA.