1. Palaeobotany Group, Institute of Geology and Palaeontology, University of Münster, Münster 48149, Germany.
2. Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China.
3. School of Ocean and Earth Science, University of Southampton, National Oceanography Centre, Southampton SO14 3ZH, UK.
4. Division of Agricultural and Environmental Sciences, School of Biosciences, The University of Nottingham, Sutton Bonington Campus, Sutton Bonington, Leicestershire LE12 5RD, UK.
5. Geography, School of Social Sciences, Oxford Brookes University, Oxford OX3 0BP, UK.
6. State Key Laboratory of Palaeobiology and Stratigraphy and Center for Excellence in Life and Paleoenvironment, Nanjing 210008, China.